Barros F (2001) Ghost crabs as a tool for rapid assessment of human impacts on exposed sandy beaches. Biol Conserv. 97: 399-404.

Bellwood O (2002) The occurrence, mechanics and significance of burying behavior in crabs (Crustacea: Brachyura). J Nat Hist. 36: 1223-1238.

Bliss DE and Mantel LH (1968) Adaptations of crustaceans to land: a summary and analysis of new findings. Am Zool. 8: 673-685.

Casadío S, Feldmann RM, Parras A and Schweitzer CE  (2005) Miocene fossil Decapoda (Crustacea: Brachyura) from Patagonia, Argentina, and their paleoecological setting. Ann Carnegie Mus. 74: 151-188.

Farrelly CA and Greenaway P (1992) Morphology and ultrastructure of the gills of terrestrial crabs (Crustacea, Gecarcinidea and Grapsidae): Adaptations for air-breathing.  Zoomorphology. 112: 39-49.

Garassino A, De Angeli A, Pasini G and Tangocci F (2010) Ocypode italica n. sp. (Crustacea, Decapoda, Brachyura, Ocypodidae) from the Piacentian (middle Pliocene) of Tuscany (Central Italy). Atti Soc Ital Sci Nat Mus Civ Stor Nat Milano. 151: 135-144.

Gibbs PE (1978) Macrofauna of the intertidal sand flats on low wooded islands, Northern Great Barrier Reef. Philos Trans R Soc Lond B. 284: 81-97.

Greenaway P and Farrelly CA (1984) The venous system of the terrestrial crab Ocypode cordimanus (Desmarest 1825) with particular reference to the vasculature of the lungs. J Morphol. 181: 133-142.

Guinot-Dumortier D and Dumortier B (1960) La stridulation chez le crabs. Crustaceana. 1: 117-155.

Hickman Jnr CP, Roberts LS, Keen SL, Larson A, l’Anson H and Eisenhour DJ (2007) Integrated principles of zoology. 14^th ed. McGraw-Hill, New York.

Horch K (1975) The acoustic behavior of the ghost crab Ocypode cordimana Latreille, 1818 (Decapoda, Brachyura). Crustaceana. 29: 193-205.

Huang J-F, Yang S-L and Ng PKL (1998) Notes on the taxonomy and distribution of two closely related species of ghost crabs, Ocypode sinensis and O. cordimanus (Decapoda, Brachyura, Ocipodidae). Crustaceana. 71: 942-954.

IUCN 2011 (2011) The IUCN red list of threatened species. International Union of Conservation of Nature. 

Jones DA (1972) Aspects of the ecology and behaviour of Ocypode ceratophthalmus (Pallas) and O. kublii de Haan (Crustacea: Ocypodidae). J Exp Mar Biol Ecol. 8: 31-43.

Jones DS and Morgan GJ (1994) A field guide to crustaceans of Australian waters.  Reed Books, Chatswood, NSW,  Australia.

Jones, D and Morgan, G (2002). A field guide to crustaceans of Australian waters. Western Australian Museum/Reed New Holland Publication, Sydney.

Jones AR, Gladstone W and Hacking NJ (2007) Australian sandy-beach ecosystems and climate change: ecology and management. Aust Zool. 34: 190-202.

Jones DS (1988) The occurrence of Ocypode pallidula  Jacquinot (Decapoda, Brachyura) in Australia and the Coral Sea. Crustaceana. 54: 33-38.

Lewinsohn CH (1977) Die Ocypodidae des Roten Meeres (Crustacea Decapoda, Brachyura) Zool Verh. 152: 45-84.

Moss D and McPhee DP (2006) The impacts of recreational four-wheel driving on the abundance of the ghost crab (Ocypode cordimanus) on subtropical beaches in SE Queensland. Coast Manage. 34, 133–140.

Noriega R and Smeuninx R (2008) Ghost crab (Ocypode cordimanus) as a potential indicator of human impact along Gold Coast beaches. Griffith Centre for Coastal Management Research Report No. 82.

Palmer JD (1971) Comparative studies of circadian locomotory rhythms in four species of terrestrial crabs. American Midland Naturalist. 85: 97-107.

Popper AN, Salmon M and Horch KW (2001) Acoustic detection and communication by decapod crustaceans. J Comp Physiol A. 187: 83-89.

Portell RW, Turner RL and Beerensson JL (2003) Occurrence of the Atlantic ghost crab Ocypode quadrata from the upper Pleistocene to Holocene Anastasia formation of Florida. J Crustacean Biol. 23: 712-722.

Rao KR (1968) The pericardial sacs of Ocypode in relation to the conservation of water, molting, and behaviour. Am Zool. 8: 561-567. 

Rosbash M (2009) The implications of multiple circadian clock origins. PLoS Biol 7: e62.

Ruppert EE, Fox RS and Barnes RD (2004) Invertebrate Zoology.  Thomson Learning. Belmont California.

Salmon M and Horch KW (1972) Acoustic signalling and detection by  semiterrestrial crabs of the family Ocypodidae. In: Winn HE and Olla BW, eds, Behaviour of marine mammals. Plenum New York, pp. 60-96.

Schlacher TA, Dugan J, Shoeman DS, Lastra M, Jones A, Scapini F, McLachlan A and Defeo A (2007) Sandy beaches at the brink. Divers Distrib. 13: 556-560.

Schlacher TA, de Jager R and Nielsen T (2011) Vegetation and ghost crabs in coastal dunes as indicators of putative stressors from tourism. Ecol Indic. 11: 284-294.

Strachan PH, Smith RC, Hamilton DA, Taylor AC and Atkinson RJ (1999) Studies on the ecology and behaviour of the ghost crab, Ocypode cursor (L.) in northern Cyprus. Sci Mar. 63: 51-60.

Strauss J and Dircksen H (2010) Circadian clocks in crustaceans: identified neuronal and cellular systems. Front Biosci. 15: 040-1074.

Türkay M, Sakai K and Apel M (1996) The Ocypode ghost crabs (Crustacea: Decapoda: Brachyura) of the Arabian Peninsula and adjacent regions. Fauna Saudi Arabia. 15: 99-117.

Valero-Pacheco E, Alvarez F, Arbarca-Arenas L and Escobar M (2007) Population density and activity pattern of the ghost crab, Ocypode quadrata, in Veracruz, Mexico. Crustaceana. 80: 313-325.

Weinstein RB and Full RJ (1998) Performance limits of low-temperature, continuous locomotion are exceeded when locomotion is intermittent in the ghost crab. Physiol Zool. 71: 274-284.

Williams AB (1984) Shrimps, lobsters and crabs of the Atlantic coast of the eastern United States, Maine to Florida, Smithsonian Institution Press, Washington DC 550pp.

Wolcott TG (1978) Ecological role of ghost crabs, Ocypode quadrata (Fab.) on an ocean beach: scavengers or predators? J Exp Mar Biol Ecol. 31: 67–82.

Wolcott TG and Wolcott DL (1984) Impact of off-road vehicles on macroinvertebrates
of a Mid-Atlantic beach. Biol Conserv. 29: 217–240.